Short CV/Education and training

  • 1983
    Graduated from Riihiketo High School, Pori, Finland

  • 1987
    Bachelor of Science, University of Turku, Finland

  • 1989
    Master of Science, University of Turku

  • 1998
    Ph.D., University of Turku

  • 1996 – 1997
    BioPD – training program for business activities in the biotechnology field (20 credit units, including a 4-week period at the University of California San Diego, USA)

  • 1989 – 1993
    Research Assistant, Department of Genetics, University of Turku

  • 1994 – 1998
    Research Scientist, Department of Medical Biochemistry and Molecular Biology, University of Oulu

  • 1998 – 1999
    Postdoctoral Fellow, Department of Medical Biochemistry and Molecular Biology, University of Oulu

  • 1999 – 2002
    Postdoctoral Researcher, Academy of Finland

  • since 2000
    Docent of Molecular Biology, Department of Medical Biochemistry and Molecular Biology, University of Oulu, Finland

  • 2000 – 2001
    Co-leader of the Enzyme Group, Collagen Research Unit, University of Oulu

  • 2001 – 2006
    Senior Research Associate, Department of Medical Biochemistry and Molecular Biology, University of Oulu

  • since 2002
    Leader of the Enzyme Group, Collagen Research Unit, University of Oulu

  • 2002 – 2007
    Academy Research Fellow, Academy of Finland

  • since 2003
    Vice Director, Collagen Research Unit (since 2008 Oulu Center for Cell-Matrix Research), Academy of Finland Center of Excellence 2000-2005, University of Oulu

  • since 2004
    Project Leader, Biocenter Oulu

  • 2007
    Vice Director, Department of Medical Biochemistry and Molecular Biology, University of Oulu

  • since 2007
    Professor of Molecular Biology, Department of Medical Biochemistry and Molecular Biology, University of Oulu

  • 2008 – 2009
    1st Vice Director (Research), Institute of Biomedicine, University of Oulu

  • since 2010
    Scientific Director, Biocenter Oulu, University of Oulu

  • 2010 – 2012
    Head, Department of Medical Biochemistry and Molecular Biology, University of Oulu

  • 2012 – 2017
    Deputy Director, Academy of Finland Center of Excellence in Cell-Extracellular Matrix Research

Selected publications

  • Published about 90 original articles or reviews in peer-reviewed international scientific journals (congress abstracts are not included in this number)



Complete list of publications

Selected projects

  • Finnish Center of Excellence in Cell-Extracellular Matrix Research

  • Mechanistic aspects of the hypoxia response and its key enzymes

  • The prolyl 4-hydroxylases that regulate the hypoxia response and their individual roles in the control of oxygen homeostasis

Membership in scientific bodies/juries

  • Finnish Society for Geneticists in Health Care, since 1990

  • Finnish Connective Tissue Research Society, since 1994; Secretary, 2000-2001, Vice Chair, 2004-2005, Chair, 2006-2007

  • International Society for Matrix Biology, since 1999; Member of the Council, since 2006

  • Societas Biochemica, Biophysica et Microbiologica Fenniae, since 2001

  • American Society for Biochemistry and Molecular Biology, 2002

  • Member of the Board, Department of Medical Biochemistry and Molecular Biology, University of Oulu, since 2007

  • Member of the Working Group of the Model Organisms Infrastructure Network, Biocenter Finland 2007 – 2008

  • Member of the Nordic Network of Caenorhabditis elegans (funded by Nordforsk), 2007 – 2010

  • Member of the Oulu Board of Directors of the TEKES programme TULI, (From Research To Busines), 2008

    Research To Business)

  • Member of the Board, Institute of Biomedicine, University of Oulu, since 2008

  • Member of the Research Committee, Medical Faculty, University of Oulu, since 2008

  • Member of the Management Committee and Working Group Leader, COST Action, TD0901, 2009-2013: Hypoxia sensing, signalling and adaptation

  • Member of the Research Council, University of Oulu, since 2010

  • Member of the Faculty Council, Medical Faculty, University of Oulu, since 2010

  • Member of the Board, Biocenter Finland, since 2010

  • Member of the Research Council for Health, Academy of Finland, 2010 – 2015; 2nd Vice Chair 2010 – 2012; 1st Vice Chair 2013 – 2015

  • Member and Vice Chair of the Subcommittee for International Joint Projects, Academy of Finland, 2010 – 2012

  • Scientific Secretary of an Expert Panel of the Research Council of Health, Academy of Finland to assess appropriations for

    • Research projects, 1999

    • Senior scientist, Senior fellow and Postdoctoral researcher posts, 2000

  • Member of the Editorial Board, Journal of Biological Chemistry, 2008 – 2012

  • Ad hoc referee for several scientific journals (Nature, Science, Cancer Research, Bone, Biochemical Journal, Structure, Matrix Biology, Cell and Tissue Research, FEBS Letters, Trends in Molecular Medicine, Cellular and Molecular Life Sciences, Analytical Biochemistry, Cancer Letters, ChemBioChem, Annals of Medicine, Clinical and Experimental Metastasis, Journal of Biotechnology, Biotechnology Progress, Biochimica et Biophysica Acta, Cellular and Molecular Biochemistry)

  • Collaborating Scientist, FibroGen Inc., South San Francisco, CA, USA, since 2010

Media coverage

  • Center for Scientific Computing, Finland, Funet-TV: Koossa pitävä voima, a film of the Center of Excellence in Molecular Biology and Pathology of Collagens and Enzymes of Collagen Biosynthesis, 2002

  • Interview to the Journal Bioteknologia Info of Tekes, the Finnish Funding Agency for Technology, 2003

  • Interview to the University of Oulu Journal Aktuumi, 2006

  • Interview to Kaleva, 2007

  • Interview to Genetic Engineering and Biotechnology News, 2007

  • Interview to Radio Oulu, 2007

  • Interview to Lääkärisanomat, 2008

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