Short CV/Education and training

  • 1989 – 1997
    Studied biology at the University of Cologne and at the Freie Universität (FU) Berlin, Germany

  • 1999 – 2004
    Doctoral student at the Freie Universität (FU) Berlin

  • 2004 – 2005
    Postdoc at the Freie Universität (FU) Berlin

  • 2005 – 2006
    Postdoc at the German Primate Centre (DPZ)

  • 2005
    Postdoc at the University of Chicago, IL, USA

  • 2006 – 2007
    Postdoc at the Bogor Agricultural University, Indonesia

  • Since 2006
    Head of the Macaca Nigra Project (

  • 2007 – 2009
    Postdoc at the German Primate Centre

  • Since 2008
    Taught in the Departments of Anthropology and Biology, University of Göttingen, Germany

  • 2009 – 2015
    Member of the Courant Research Centre "Evolution of Social Behaviour", University of Göttingen

  • 2009 – 2015
    Head of the Emmy Noether junior research group "Sexual Selection among Primates" at the German Primate Centre (DPZ)

  • Since 2016
    Reader (Associate Professor) for Primate Behaviour and Conservation at the Liverpool John Moores University, UK

Selected publications

  • Engelhardt, A. et al.: Information content of female copulation calls in wild long-tailed macaques (Macaca fascicularis). Behavioral Ecology and Sociobiology 66 pp. 121-134.

  • Higham, J. et al.: Colour signal information content and the eye of the beholder: a case study in the rhesus macaque. In: Behavioral Ecology 21. pp. 739-746.

  • Neumann, C. et al.: Loud calls in male Macaca nigra – a signal of dominance in a tolerant primate species. In: Animal Behaviour 79, 2010. pp. 187-193.

  • Girard-Butoz, C. et al.: Seasonal and social influences on fecal androgen and glucocorticoid excretion in wild male long-tailed macaques (Macaca fascicularis). In: Physiology and Behavior 98, 2009. pp. 168-175.

  • Brauch, K. et al.: Paternity in Barbary macaques (Macaca sylvanus): both, males and females pull the strings. In: Behavioral Ecology and Sociobiology 62, 2008. pp. 1453-1466.

  • Engelhardt, A., Hodges, J.K., Heistermann, M.: Post-conception mating in wild long-tailed macaques (Macaca fascicularis): characterization, endocrine correlates and functional significance. In: Hormones and Behavior 51, 2007. pp. 3-10.

  • Pradhan, G. et al.: The evolution of female copulation calls in primates: a review and a new model. In: Behavioral Ecology and Sociobiology 59, 2006. pp. 333-343.

  • Engelhardt, A. et al.: Determinants of male reproductive success in wild long-tailed macaques (Macaca fascicularis) – male monopolisation, female mate choice or post-copulatory mechanisms? In: Behavioral Ecology and Sociobiology 59, 2006. pp. 740-752.

  • Engelhardt, A. et al.: Female sexual behavior, but not sex skin swelling reliably indicates the timing of the fertile phase in wild long-tailed macaques (Macaca fascicularis). In: Hormones and Behavior 47, 2005. pp. 195-204.

  • Engelhardt, A. et al.: Assessment of female reproductive status by male long-tailed macaques (Macaca fascicularis) under natural conditions. In: Animal Behaviour 67, 2004. pp. 915-924.

Complete list of publications

Selected projects

  • Determinants of male reproductive success in multi-male groups

  • Function of sexual signals

  • Hormone-behaviour relationships

  • Sperm competition in macaques

  • Communication and cognition

Membership in scientific bodies/juries

  • Spokesperson for the section Behavioural Biology in the Deutsche Zoologische Gesellschaft (German Zoological Association)

  • Research proposal reviewer for the German Research Foundation (DFG)

  • Reviewed applicants for the International Primatological Society (Research Grants)

  • Reviewed applicants for the Leakey Foundation

Additional qualifications

  • Head of an Emmy Noether junior research group, funded by the German Research Foundation (DFG)

Soft Skills/Other activities and achievements

Other activities and achievements/family

  • Three children


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