Short CV/Education and training

  • Since 2014
    Margarete von Wrangell-Fellow

  • Since 2014
    Head of Junior Research Project Group NeuroLearn, Leibniz-Institut fuer Wissensmedien Tuebingen

  • 2012
    Doctorate (PhD), Tuebingen University

  • 2011
    Equivalent to MSc in Psychology, University of Tuebingen

  • 2009
    Doctorate (MD), RWTH Aachen University

  • 2008 – 2012
    Section of Diagnostics and Cognitive Neuropsychology, Institute of Psychology, University of Tuebingen

  • 2008 – 2012
    Department of Neurology, Section Neuropsychology and Section Neurolinguistic, University Hospital, RWTH Aachen University

  • 2007
    Medical Approbation and Graduation from Medical School

Selected publications

  • Bloechle, J., Huber, S., Bahnmueller, J., Rennig, J., Willmes, K., Cavdaroglu, S., Moeller, K., & Klein, E. (2016). Fact Learning in Complex Arithmetic – The Role of the Angular Gyrus revisited. Human Brain Mapping, 37, 3061-3079.

  • Klein, E., Suchan, J., Moeller, K., Karnath, H.-O., Knops, A., Wood, G., Nuerk, H.-C., & Willmes K. (2016). Considering structural connectivity in the triple code model of numerical cognition – Differential connectivity for magnitude processing and arithmetic facts. Brain Structure and Function, 221(2), 979-95.

  • Jung, S., Halm, K., Huber, W., Willmes, K., & Klein, E. (2015). What letters can "learn" from Arabic digits – fMRI-controlled single case therapy study of peripheral agraphia. Brain and Language, 149, 13-26.

  • Klein, E., Huber, S., Nuerk, H.-C., & Moeller, K. (2014). Operational momentum affects eye fixation behaviour. Quarterly Journal of Experimental Psychology, 67(8), 1614-1625.

  • Mihulowicz, U., Willmes, K., Karnath, H.-O., & Klein, E. (2014). Single-digit arithmetic processing – anatomical evidence from statistical voxel-based lesion analysis. Frontiers in Human Neuroscience, 8:286.

  • Klein, E., Mann, A., Huber, S., Bloechle, J., Willmes, K., Karim, A.A., Nuerk, H.-C., & Moeller, K. (2013). Bilateral bi-cephalic tDCS with two active electrodes of the same polarity modulates bilateral cognitive processes differentially. PLoS ONE 8(8): e71607.

  • Klein, E., Moeller, K., Glauche, V., Weiller, C., & Willmes, K. (2013). Processing pathways in mental arithmetic – Evidence from probabilistic fiber tracking. PLoS ONE, 8(1): e55455.

  • Klein, E., Moeller, K., Willmes, K., Nuerk, H.-C., & Domahs, F. (2011). The influence of implicit hand-based representations on mental arithmetic. Frontiers in Psychology, 2, 197.

Complete list of publications

Selected projects

  • Neurocognitive foundations and processing pathways of arithmetic learning and their plasticity in adults as well as in typically and atypically developing children (

  • Benefits of a game-based cognitive interface for knowledge work – from basic effects and neural correlates to neuropsychological rehabilitation (


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