Short CV/Education and training


  • Since 2003
    Head of Section for Genetics and Bioinformatics

  • 1997
    Professor of Animal Genetics, Division of Animal Breeding and Genetics, RVAU

  • 1991
    Associate professor, Department of Animal Genetics, RVAU

  • 1988
    Assistant professor, Department of Animal Genetics, RVAU

  • 1986
    Post doctoral fellowship at Research Institute of Scripps Clinic, San Diego, USA

  • 1984
    Assistant professor, Department of Animal Genetics, RVAU

  • 1982
    Research assistant, Department of Animal Genetics, RVAU


  • 1996

  • 1986
    Ph.D. degree in Animal Genetics

  • 1982
    DVM, The Royal Veterinary and Agricultural University, (RVAU), Copenhagen, Denmark

Selected publications

  • Gorodkin, J., Cirera, S., Gilchrist, M., Paniz, F., Jørgensen, C.B., Scheiby-Knudsen, K., Arvin, T., Lumholt, S., Sawera, M., Green, T., Nielsen, B.J., Havgaard, J.H., Stærfelt, H.H., Brunak, Rosenkilde, C., S., Madsen, L.B., Thomsen, B., Hornshøj, H., Zhan, B., Wang, X., Yang, H., Bendixen, C and Fredholm M. 2007. Porcine transcriptome analysis based on 97 non-normalized cDNA libraries and assembly of 1,021,891 ESTs. Genome Biology 8 (4):R45.

  • Charlier C., Coppieters W., Rollin F., Desmecht D., Agerholm J.S., Cambisano N., Carta E., Dardano S., Dive M., Fasquelle C., Frennet J.C., Hanset R., Hubin X., Jorgensen C.B., Karim L., Kent M., Harvey K., Pearce B.R., Simon P., Tama N., Nie H., Vandeputte S., Lien S., Longeri M., Fredholm M., Harvey R.J. and Michel Georges. 2008. Highly effective SNP-based association mapping and management of recessive defects in livestock. Nature Genetics 40: 449-454.

  • Bruun CS, Jorgensen CB, Bay L, Cirera S, Jensen HE, Leifsson PS, Nielsen J, Christensen K and Fredholm M. 2008. Phenotypic and genetic characterization of a novel phenotype in pigs characterized by juvenile hairlessness and age dependent emphysema. BMC Genomics, 9:283 doi:10.1186/1471-2164-9-283

  • Nygaard AB, Cirera S, Gilchrist MJ, Gorodkin J, Jørgensen CB and Fredholm M. A study of alternative splicing in the pig 2010. BMC Research Notes 3:123.

  • Podolska A, Anton C, Bak M, Tommerup N, Skovgaard K, Heegaard PM, Gorodkin J, Cirera S & Fredholm M. Profiling microRNAs in lung tissue from pigs infected with Actinobacillus pleuropneumoniae. BMC Genomics 2012 Sep 6,13:459.

  • Larson G, Karlsson EK, Perri A, Webster MT, Ho SYW, Peters J, Stahl PW, Piper PJ, Lingaas F, Fredholm M, Comstock KE, Modiano JF, Schelling C, Agoulnik AI, Leegwater PA, Dobney K, Vigne J-D, Vila C, Andersson L, Lindblad-Toh K. 2012. Rethinking dog domestication be integrating genetics, archeology, and biogeography. PNAS. 109(23): 8878-8883

  • Rubin C-J, Megens H-J, Barrio AM, Maqbool K, Sayyab S, Schwochow D, Wang C, Carlborg Ö, Jern P, Jørgensen CB, Archibald AL, Fredholm M, Groenen MAM and Andersson L. 2012. Strong signatures of selection in the domestic pig genome. PNAS, doi:10.1073/pnas.1217149109.

  • Bruun CS, Jäderlund KH, Berendt M, Jensen KB, Spodsberg EH, Gredal H, Shelton GD, Mickelson JR, Minor KM, Lohi H, Bjerkås I, Stigen Ø, Espenes A, Rohdin C, Edlund R, Ohlsson J, Cizinauskas S, Leifsson PS, Drögemüller C, Moe L, Cirera S and Fredholm M. 2013. A Gly98Val mutation in the N-Myc Downstream Regulated Gene 1 (NDRG1) in Alaskan Malamutes with Polyneuropathy. PLoS One: 8(2):e54547. doi: 10.1371.

  • Mentzel CMJ, Anthon C, Jacobsen MJ, Karlskov-Mortensen P, Bruun CS, Jørgensen CB , Gorodkin J, Cirera S and Fredholm M. Gender and obesity specific microRNA expression in adipose tissue from lean and obese pigs. PLOS ONE | DOI:10.1371/journal.pone.0131650 July 29, 2015.

  • Pant SD, Karlskov-Mortensen P, Jacobsen MJ, Cirera S, Kogelman LJA, Bruun CS, Mark T, Jørgensen CB, Grarup N, Appel EVR, Galjatovic EAA, Hansen T, Pedersen O, Guerin M, Huby T, Lesnik P, Meuwissen THE, Kadarmideen HN and Fredholm M. Comparative Analyses of QTLs influencing Obesity and Metabolic Phenotypes in Pigs and Humans. PlosOne, 2015 Sep 8;10(9):e0137356. doi: 10.1371/journal.pone.0137356.

Selected projects

  • Cutting edge genomics for sustainable breeding

  • Characterization of a pig model for chronic obstructive lung disease

  • Unravelling the molecular basis of common complex human disorders using the dog as a model

  • Genetic studies of obesity and obesity related metabolic diseases in a pig model

  • MetaPig

Membership in scientific bodies/juries

  • Member of International Society of Animal Genetics (ISAG)

  • Member of the Danish Academy of Technical Sciences

Editorial boards and referee

  • Animal Genetics; Molecular Genetics and Genomics, Canine Genetics and Epidemiology, Journal of Applied Genetics; Animal Biotechnology; BMC Genomics; BMC Genetics, PLoS Genetics; USDA competitive grants; Biotechnology programs, Norwegian Agricultural and Veterinary Research Council; Innovation Foundation; Biotechnology and Biological Sciences Research Council, UK; INTAS


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